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LIPN Gene

protein-coding GIFtS: 33

GC10P084155

lipase, family member N
(Previous names: lipase-like, ab-hydrolase domain containing 4 )
Symbol approved by the HUGO Gene Nomenclature Committee (HGNC) database
(Previous symbol: LIPL4)

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(According to ¹HGNC, ²Entrez Gene,
³UniProtKB/Swiss-Prot, ⁴UniProtKB/TrEMBL, ⁵OMIM, ⁶GeneLoc , and/or ⁷Ensembl, ⁸miRBase)
About This Section

Aliases & Descriptions
lipase, family member N¹ ²		LIPL4² ³
lipase-like, ab-hydrolase domain containing 4¹ ²		lipase N²
bA186O14.3¹ ²		lipase member N²
Lipase-like abhydrolase domain-containing protein 4² ³		EC 3.1.1.-³

External Ids:	HGNC: 23452¹	Entrez Gene: 643418²	Ensembl: ENSG00000204020⁷	UniProtKB: Q5VXI9³

Export aliases for LIPN gene to outside databases

Previous GC identifer: GC10P090508

(According to Entrez Gene, Tocris Bioscience, Wikipedia's Gene Wiki,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

UniProtKB/Swiss-Prot: LIPN_HUMAN, Q5VXI9

Function: Plays a highly specific role in the last step of keratinocyte differentiation. May have an essential function

in lipid metabolism of the most differentiated epidermal layers

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to UCSC (hg19) and Ensembl (release 60), Regulatory elements and Epigenetics data according to Qiagen and/or SABiosciences)
About This Section

Regulatory elements:

Search SABiosciences Regulatory transcription factor binding sites for LIPN
Other transcription factors

Search SABiosciences Chromatin IP Primers for LIPN

Epigenetics:

QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays

Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track

Entrez Gene cytogenetic band: 10q23.31 Ensembl cytogenetic band: 10q23.31 HGNC chromosome: 10

LIPN Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)
LIPN gene location

GeneLoc gene densities for chromosome 10 GeneLoc Exon Structure

GeneLoc location for GC10P084155: view genomic region (about GC identifiers)

Start:	84,155,904 bp from pter
End:	84,172,685 bp from pter
Size:	16,782 bases
Orientation:	plus strand

1 alternative location:

Chr10+ 90,519,952-90,537,999

RefSeq DNA sequence:

NC_000010.10 NT_030059.13

(According to ¹UniProtKB, neXtProt, and/or Ensembl, Phosphorylation sites according to ²Phosphosite, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from Millipore, Sigma-Aldrich, R&D Systems, GenScript, Enzo Life Sciences, OriGene, Novus Biologicals, Sino Biological, and/or ProSpec,
Biochemical Assays by Millipore, Sigma-Aldrich, R&D Systems, OriGene, GenScript, Cell Signaling Technology, Enzo Life Sciences, and/or Uscn, Ontologies according to Gene Ontology Consortium 01 Dec 2010 and Entrez Gene, Antibodies by Millipore, Sigma-Aldrich, R&D Systems, GenScript, Cell Signaling Technology, OriGene, Novus Biologicals, and/or Epitomics)
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UniProtKB/Swiss-Prot: LIPN_HUMAN, Q5VXI9 (See protein sequence)

Recommended Name: Lipase member N precursor

Size: 398 amino acids; 45534 Da

Subcellular location: Secreted (Potential)

Secondary accessions: A7KIH9

Explore the universe of human proteins at neXtProt for LIPN: NX_Q5VXI9

REFSEQ proteins: NP_001095939.1

ENSEMBL proteins:

ENSP00000383923

Human Recombinant Proteins

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1 Gene Ontology (GO) cellular component term (GO ID links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0005576	extracellular region	IEA	--

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LIPN for ontologies About GeneDecksing

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(According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
About This Section

LIPN for domains About GeneDecksing

2 InterPro domains/families:

IPR006693 AB_hydro-assoc_lipase

IPR000073 AB_hydrolase_1

Graphical View of Domain Structure for InterPro Entry Q5VXI9

ProtoNet protein and cluster: Q5VXI9

UniProtKB/Swiss-Prot: LIPN_HUMAN, Q5VXI9

Similarity: Belongs to the AB hydrolase superfamily. Lipase family

(According to UniProtKB, IUBMB,and/or Genatlas, Animal models from MGI Dec 24 2010,
shRNA from OriGene, Sigma-Aldrich, RNAi from Millipore, siRNAs from Sigma-Aldrich, OriGene, Qiagen, Super-pooled esiRNAs from Sigma-Aldrich, microRNA from Sigma-Aldrich, Qiagen, SABiosciences, Clones from Millipore, Sigma-Aldrich, OriGene, GenScript, Sino Biological, Cell Lines from GenScript, Ontologies according to Gene Ontology Consortium 01 Dec 2010 via Entrez Gene.)
About This Section

UniProtKB/Swiss-Prot: LIPN_HUMAN, Q5VXI9

Function: Plays a highly specific role in the last step of keratinocyte differentiation. May have an essential function

in lipid metabolism of the most differentiated epidermal layers

Enzyme Number (IUBMB): EC 3.1.1.-

Inhib. RNA:		Browse for Gene Knock-down Tools from Millipore
		Browse Nano Scale siRNA at Sigma-Aldrich Sigma-Aldrich esiRNA and siRNA for LIPN Sigma-Aldrich shRNA for LIPN
		OriGene 29mer shRNA kits in GFP-retroviral vector (see all 3): LIPN OriGene shRNA RFP (see all 3): LIPN OriGene basic RS shRNA (see all 3): LIPN OriGene siRNA (see all 3): LIPN
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Gene
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Cell
Lines:

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1 Gene Ontology (GO) molecular function term (GO ID links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0016787	hydrolase activity	IEA	--

About this table

LIPN for ontologies About GeneDecksing

(Pathways according to Millipore, Cell Signaling Technology, Sigma-Aldrich, KEGG and/or UniProtKB, Sets of similar genes according to GeneDecks, PCR Arrays from SABiosciences, Proteins Network according to SABiosciences, Sigma-Aldrich, Interactions according to ¹UniProtKB, ²MINT, and/or ³STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Dec 2010 via Entrez Gene).
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LIPN for pathways About GeneDecksing

1 Sigma-Aldrich "Your Favorite Gene" (powered by Ingenuity) Pathway for LIPN

Xenobiotic Metabolism Signaling

SABiosciences Custom Pathway-Focused PCR Arrays for LIPN

Sigma-Aldrich "Your Favorite Gene" (powered by Ingenuity) Molecular Interaction Network for LIPN

Search SABiosciences Gene Network Central^TM Interacting Genes and Proteins Networks for LIPN

2 Gene Ontology (GO) biological process terms (GO ID links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0006629	lipid metabolic process	--	--
GO:0016042	lipid catabolic process	IEA	--

About this table

LIPN for ontologies About GeneDecksing

(Chemical Compounds according to UniProtKB, Enzo Life Sciences, Sigma-Aldrich, Tocris Bioscience, and/or Novoseek and Drugs according to Enzo Life Sciences and/or PharmGKB)
About This Section

Browse Small Molecules at Sigma-Aldrich

Browse drugs & compounds from Enzo Life Sciences

Browse Tocris compounds for LIPN

(GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 228 Homo sapiens; Dec 8 2010) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView, transcript ids from Ensembl with links to UCSC,
non coding RNAs according to RNAdb,
ESTs according to GeneTide,
exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
RNAi Products from Millipore,
siRNAs from Sigma-Aldrich, OriGene, Qiagen, Super-pooled esiRNAs from Sigma-Aldrich,
shRNA from Sigma-Aldrich, OriGene, microRNA from Sigma-Aldrich, Qiagen, SABiosciences,
Tagged/untagged cDNA clones from OriGene, Sigma-Aldrich, GenScript, Primers from OriGene and/or SABiosciences)
About This Section

Inhib. RNA:		Browse for Gene Knock-down Tools from Millipore
		Browse Nano Scale siRNA at Sigma-Aldrich Sigma-Aldrich esiRNA and siRNA for LIPN Sigma-Aldrich shRNA for LIPN
		OriGene 29mer shRNA kits in GFP-retroviral vector (see all 3): LIPN OriGene shRNA RFP (see all 3): LIPN OriGene basic RS shRNA (see all 3): LIPN OriGene siRNA (see all 3): LIPN
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miRNA:		Browse microRNA Mimics at Sigma-Aldrich Browse microRNA target validation systems at Sigma-Aldrich
		QIAGEN Custom miScript Target Protector blocks miRNA-binding site of LIPN
		Search SABiosciences for Assays for microRNAs that regulate LIPN

Clones:		Browse FLAG tag genes at Sigma-Aldrich
		OriGene GFP tagged cDNA clones in CMV expression vector: LIPN OriGene Myc/DDK tagged cDNA clones in CMV expression vector: LIPN OriGene untagged cDNA clones in CMV expression vector: LIPN Browse 3'-UTR reporter clones for miRNA target validation Browse OriGene MicroRNA Expression Plasmids
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Primers:		OriGene genome-wide validated SYBR primer pairs: LIPN Browse OriGene validated miRNA SYBR primer pairs
		SABiosciences RT² qPCR Primer Assay for LIPN: PPH63511A

REFSEQ mRNAs for LIPN gene:

NM_001102469.1

Additional cDNA sequence: EF426483.1

2 DOTS entries:

DT.121293039 DT.121268916

Unigene Cluster for LIPN:

Lipase, family member N

Hs.632091 [show with all ESTs]

Unigene Representative Sequence: EF426483

GeneLoc Exon Structure

2 Ensembl transcripts including schematic representations, and UCSC links where relevant:

ENST00000491766 ENST00000404459(uc010qmw.1)

(Experimental results according to ¹GeneNote and GNF BioGPS,
probe sets-to-genes annotations according to ²GeneAnnot , ³GeneTide , Sets of similar genes according to GeneDecks, Electronic Northern calculations according to data from UniGene (Build 228 Homo sapiens), SAGE tags according to CGAP, plus additional links to SOURCE, and/or GNF BioGPS, and/or EXPOLDB, and/or UniProtKB,
Primers from OriGene and/or SABiosciences )
About This Section

LIPN expression in normal and diseased human tissues

¹ Data from GNF BioGPS
About these images

LIPN gene electronic northern expression

About these images

CGAP SAGE TAG: ATGTCCTCGT

SOURCE GeneReport for Unigene cluster: Hs.632091

UniProtKB/Swiss-Prot: LIPN_HUMAN, Q5VXI9

Tissue specificity: Exclusively expressed in the epidermis within the granular keratinocytes

Primers:		OriGene genome-wide validated SYBR primer pairs: LIPN Browse OriGene validated miRNA SYBR primer pairs
		SABiosciences RT² qPCR Primer Assay for LIPN: PPH63511A

SABiosciences Custom PCR Arrays for LIPN

(Orthologs according to ^1,2HomoloGene (²older version, for species not in ¹newer version), ³euGenes, ⁴SGD and/or ⁵MGI Dec 24 2010, with possible further links to Flybase and/or WormBase, Gene Trees according to Ensembl)
About This Section

Orthologs for LIPN gene from 3 species

Organism	Gene	Locus	Description	Human Similarity	NCBI accessions
chimpanzee (Pan troglodytes)	LIPN¹	--	lipase, family member N	99.24(n) 98.74(a)	466144 XM_521546.2 XP_521546.2
rat (Rattus norvegicus)	Lipn¹	--	lipase, family member N	85.99(n) 82.53(a)	499345 XM_574655.2 XP_574655.2
mouse (Mus musculus)	Lipn^{1 , 5}	19 (29.32 cM)⁵	lipase, family member N^{1, 5}	84.97(n)¹ 82.12(a)¹	70166¹ NM_027340.2¹ NP_081616.1¹ AK008974⁵ AK154333⁵

About this table Species with no ortholog for LIPN

ENSEMBL Gene Tree for LIPN (if available)

(Paralogs according to ¹HomoloGene
and ²Ensembl, Pseudogenes according to ³Pseudogene.org)
About This Section

Paralogs for LIPN gene

LIPA² LIPK² LIPJ² LIPM² LIPF²

LIPN for paralogs About GeneDecksing

(SNPs according to the ¹NCBI SNP Database, ²Ensembl, ³PupaSUITE, and UniProtKB, Linkage Disequilibrium by HapMap, Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Resequencing Primers from Qiagen)
About This Section

SNP ID	Valid	Chr 10 pos	Sequence	Recs	AA Chg	Type	More	Recs	Pop	Total sample
Genomic Data				Transcription Related Data				Allele Frequencies
			--	--	--		--	--
rs188571^1,2	C,F,O,A,	90522839(-)	`ATGACC/TTCAGC`	1	--	int¹		9	NA EA WA	358
rs303461^1,2	C,F,O,A,	90519392(-)	`CAGTGC/TGTAGG`	1	--	ng5¹		18	NS NA EA	886
rs303462^1,2	C,F,A,	90520101(-)	`ttcacG/Atatgt`	1	--	ng5¹		14	NS NA EA WA	458
rs303463^1,2	C,F,A,	90520269(-)	`TTAACT/CTCTAG`	1	--	ng5¹		9	NA EA WA	358
rs303464^1,2	C,F,O,A,	90520562(-)	`ACAGCA/GTGCTC`	1	--	ng5¹		11	NA WA	228
rs303465^1,2	C,F,O,A,	90520847(-)	`ATTTGA/CCAAAA`	1	--	ng5¹		9	NA EA WA	358
rs303466^1,2	C,F,O,A,	90521616(-)	`ATGTAC/TATCAA`	1	--	int¹		13	NA NS EA WA	768
rs303467^1,2	C,F,O,A,	90533235(-)	`TTTTAT/CTTCAA`	1	--	int¹		22	NA NS EA WA	1492
rs303468^1,2	C,F,H,	90533543(-)	`TTTGAA/GCAAGC`	1	--	int¹		21	NS EA NA WA	1348
rs303469^1,2	C,F,	90533682(-)	`CTAAAA/GGCCAA`	1	--	int¹		11	NA NS EA WA	774

About this table

HapMap Linkage Disequilibrium images for LIPN (up to first 250kb)
Structural Variations (Copy Number Variations, Insertions/Deletions, Inversions)
Database of Genomic Variants (DGV) variations for LIPN: --

QIAGEN SeqTarget long-range PCR primers for resequencing LIPN

(in which this Gene is Involved, According to OMIM, UniProtKB, PharmGKB, Genatlas, GeneTests, Blood group antigen gene mutations by BGMUT, LSDB, HGMD, GAD, HuGE Navigator, and/or TGDB.)
About This Section

(Possibly Related Articles in Doctor's Guide)
About This Section

(in PubMed. Associations of this gene to articles via ¹Entrez Gene, ²UniProtKB/Swiss-Prot, ³HGNC, ⁴GAD, ⁵PharmGKB, ⁶UniProtKB/TrEMBL, and/or ⁷Novoseek)
About This Section

2 PubMed articles for LIPN gene integrated from 7 sources:
(articles sorted by number of sources associating them with LIPN)

Large-scale identification of human genes implicated in epidermal barrier function. (PubMed id 17562024)^{1, 2} Toulza E.... Guerrin M. (2007)
The DNA sequence and comparative analysis of human chromosome 10. (PubMed id 15164054)² Deloukas P.... Rogers J. (2004)

(in PubMed, OMIM, and NCBI Bookshelf)
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		AND	OR
Aliases
Free Text

Query String		PubMed OMIM NCBI Bookshelf
	(Note: In FireFox, select the above section and copy using Ctrl-C)

(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, Kegg, and/or H-InvDB)
About This Section

Entrez Gene: 643418	HGNC: 23452	Ensembl:ENSG00000204020	euGenes: HUgn643418	ECgene: LIPN
H-InvDB: LIPN

(According to HUGE)
About This Section

(According to ATLAS, HORDE, IMGT, MTDB, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
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Intellectual Property for LIPN gene
(Patent information from GeneIP, Licensable technologies from WIS Yeda, Salk, Tufts, IP news from XenneX, Inc.)

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Patent Information for LIPN gene:
Search GeneIP for patents involving LIPN

GeneCards and IP:
Japan Patent Office Licenses GeneCards European Patent Office Licenses GeneCards Improving the IP Search

(Antibodies, recombinant proteins, and assays by Millipore, Sigma-Aldrich, R&D Systems, Qiagen, GenScript, Cell Signaling Technology, SABiosciences, Novus Biologicals, Epitomics, ProSpec, Uscn,
Clones available from Millipore, Sigma-Aldrich, OriGene, GenScript, Sino Biological, PCR Arrays from SABiosciences, Drugs and/or compounds by Sigma-Aldrich, Tocris Bioscience, and/or Enzo Life Sciences)
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LIPN Gene

protein-coding GIFtS: 33

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LIPN Gene

protein-coding GIFtS: 33

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