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MARVELD1 Gene

protein-coding GIFtS: 41

GC10P093098

MARVEL domain containing 1
(Previous name: MARVEL (membrane-associating) domain containing 1 )
Symbol approved by the HUGO Gene Nomenclature Committee (HGNC) database
(Previous symbol: MRVLDC1)

		Products

(According to ¹HGNC, ²Entrez Gene,
³UniProtKB/Swiss-Prot, ⁴UniProtKB/TrEMBL, ⁵OMIM, ⁶GeneLoc , and/or ⁷Ensembl, ⁸miRBase)
About This Section

Aliases & Descriptions
MARVEL domain containing 1¹ ²		bA548K23.8¹ ²
FLJ22343¹ ²		MRVLDC1² ³
MARVEL (membrane-associating) domain containing 1¹ ²		putative MARVEL domain-containing protein 1²
MGC4415¹ ²		MARVD1²
FLJ23440¹ ²		RP11-548K23.8²
GB14¹ ²

External Ids:	HGNC: 28674¹	Entrez Gene: 83742²	Ensembl: ENSG00000155254⁷	UniProtKB: Q9BSK0³

Export aliases for MARVELD1 gene to outside databases

Previous GC identifer: GC10P099464

(According to Entrez Gene, Tocris Bioscience, Wikipedia's Gene Wiki,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to UCSC (hg19) and Ensembl (release 60), Regulatory elements and Epigenetics data according to Qiagen and/or SABiosciences)
About This Section

Regulatory elements:

SABiosciences Regulatory transcription factor binding sites in the MARVELD1 gene promoter:
         deltaCREB   CREB   ARP-1   RP58   Pax-2   MyoD   Tal-1   E47   GATA-2   GATA-1
         Other transcription factors

Search SABiosciences Chromatin IP Primers for MARVELD1

Epigenetics:

QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays for MARVELD1

Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track

Entrez Gene cytogenetic band: 10q24.2 Ensembl cytogenetic band: 10q24.2 HGNC cytogenetic band: 10q24.2

MARVELD1 Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)
MARVELD1 gene location

GeneLoc gene densities for chromosome 10 GeneLoc Exon Structure

GeneLoc location for GC10P093098: view genomic region (about GC identifiers)

Start:	93,098,658 bp from pter
End:	93,103,102 bp from pter
Size:	4,445 bases
Orientation:	plus strand

1 alternative location:

Chr10+ 99,472,930-99,484,694

RefSeq DNA sequence:

NC_000010.10 NT_030059.13

(According to ¹UniProtKB, neXtProt, and/or Ensembl, Phosphorylation sites according to ²Phosphosite, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from Millipore, Sigma-Aldrich, R&D Systems, GenScript, Enzo Life Sciences, OriGene, Novus Biologicals, Sino Biological, and/or ProSpec,
Biochemical Assays by Millipore, Sigma-Aldrich, R&D Systems, OriGene, GenScript, Cell Signaling Technology, Enzo Life Sciences, and/or Uscn, Ontologies according to Gene Ontology Consortium 01 Dec 2010 and Entrez Gene, Antibodies by Millipore, Sigma-Aldrich, R&D Systems, GenScript, Cell Signaling Technology, OriGene, Novus Biologicals, and/or Epitomics)
About This Section

UniProtKB/Swiss-Prot: MALD1_HUMAN, Q9BSK0 (See protein sequence)

Recommended Name: Putative MARVEL domain-containing protein 1

Size: 173 amino acids; 18914 Da

Subcellular location: Membrane; Multi-pass membrane protein (Potential). Nucleus. Note=According to PubMed:19364627,

located in nucleus instead of cell membrane

Miscellaneous: Down-regulated in primary multiple tumors derived from ovary, vulva, uterus, cervix, breast, testis,

kidney bladder and liver. The reduced expression is owing to DNA methylation and could be reversed by pharmacologic

demethylation

Caution: Product of a dubious CDS prediction. May be produced at very low levels due to a premature stop codon in the

mRNA, leading to nonsense-mediated mRNA decay

Explore the universe of human proteins at neXtProt for MARVELD1: NX_Q9BSK0

REFSEQ proteins: NP_113672.1
Human Recombinant Proteins

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3 Gene Ontology (GO) cellular component terms (GO ID links to tree view):

GO ID	Qualified GO term	Evidence	PubMed IDs
GO:0005634	nucleus	IEA	--
GO:0016020	membrane	IEA	--
GO:0016021	integral to membrane	IEA	--

About this table

MARVELD1 for ontologies About GeneDecksing

Antibodies for MARVELD1:

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Assays for MARVELD1:

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	Browse ELISAs and CLIAs at Uscn

(According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
About This Section

MARVELD1 for domains About GeneDecksing

2 InterPro domains/families:

IPR021128 MARVEL-like_dom

IPR008253 Marvel

Graphical View of Domain Structure for InterPro Entry Q9BSK0

ProtoNet protein and cluster: Q9BSK0

2 Blocks protein families:

IPB000408 Regulator of chromosome condensation

IPB008253 Marvel

UniProtKB/Swiss-Prot: MALD1_HUMAN, Q9BSK0

Similarity: Contains 1 MARVEL domain

(According to UniProtKB, IUBMB,and/or Genatlas, Animal models from MGI Dec 24 2010,
shRNA from OriGene, Sigma-Aldrich, RNAi from Millipore, siRNAs from Sigma-Aldrich, OriGene, Qiagen, Super-pooled esiRNAs from Sigma-Aldrich, microRNA from Sigma-Aldrich, Qiagen, SABiosciences, Clones from Millipore, Sigma-Aldrich, OriGene, GenScript, Sino Biological, Cell Lines from GenScript, Ontologies according to Gene Ontology Consortium 01 Dec 2010 via Entrez Gene.)
About This Section

Inhib. RNA:		Browse for Gene Knock-down Tools from Millipore
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		OriGene 29mer shRNA kits in GFP-retroviral vector (see all 2): MARVELD1 OriGene shRNA RFP (see all 2): MARVELD1 OriGene basic RS shRNA (see all 2): MARVELD1 Browse OriGene siRNA
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miRNA:		Browse microRNA Mimics at Sigma-Aldrich Browse microRNA target validation systems at Sigma-Aldrich
		QIAGEN Custom miScript Target Protector blocks miRNA-binding site of MARVELD1
		Search SABiosciences for Assays for microRNAs that regulate MARVELD1

Gene
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Cell
Lines:

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(Pathways according to Millipore, Cell Signaling Technology, Sigma-Aldrich, KEGG and/or UniProtKB, Sets of similar genes according to GeneDecks, PCR Arrays from SABiosciences, Proteins Network according to SABiosciences, Sigma-Aldrich, Interactions according to ¹UniProtKB, ²MINT, and/or ³STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Dec 2010 via Entrez Gene).
About This Section

SABiosciences Custom Pathway-Focused PCR Arrays for MARVELD1

Search SABiosciences Gene Network Central^TM Interacting Genes and Proteins Networks for MARVELD1

(Chemical Compounds according to UniProtKB, Enzo Life Sciences, Sigma-Aldrich, Tocris Bioscience, and/or Novoseek and Drugs according to Enzo Life Sciences and/or PharmGKB)
About This Section

Browse Small Molecules at Sigma-Aldrich

Browse drugs & compounds from Enzo Life Sciences

Browse Tocris compounds for MARVELD1

(GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 228 Homo sapiens; Dec 8 2010) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView, transcript ids from Ensembl with links to UCSC,
non coding RNAs according to RNAdb,
ESTs according to GeneTide,
exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
RNAi Products from Millipore,
siRNAs from Sigma-Aldrich, OriGene, Qiagen, Super-pooled esiRNAs from Sigma-Aldrich,
shRNA from Sigma-Aldrich, OriGene, microRNA from Sigma-Aldrich, Qiagen, SABiosciences,
Tagged/untagged cDNA clones from OriGene, Sigma-Aldrich, GenScript, Primers from OriGene and/or SABiosciences)
About This Section

Inhib. RNA:		Browse for Gene Knock-down Tools from Millipore
		Browse Nano Scale siRNA at Sigma-Aldrich Sigma-Aldrich siRNA for MARVELD1 Sigma-Aldrich shRNA for MARVELD1
		OriGene 29mer shRNA kits in GFP-retroviral vector (see all 2): MARVELD1 OriGene shRNA RFP (see all 2): MARVELD1 OriGene basic RS shRNA (see all 2): MARVELD1 Browse OriGene siRNA
		QIAGEN FlexiTube/FlexiPlate siRNA for gene silencing of MARVELD1

miRNA:		Browse microRNA Mimics at Sigma-Aldrich Browse microRNA target validation systems at Sigma-Aldrich
		QIAGEN Custom miScript Target Protector blocks miRNA-binding site of MARVELD1
		Search SABiosciences for Assays for microRNAs that regulate MARVELD1

Clones:		Browse FLAG tag genes at Sigma-Aldrich
		OriGene GFP tagged cDNA clones in CMV expression vector: MARVELD1 OriGene Myc/DDK tagged cDNA clones in CMV expression vector: MARVELD1 OriGene untagged cDNA clones in CMV expression vector: MARVELD1 Browse 3'-UTR reporter clones for miRNA target validation Browse OriGene MicroRNA Expression Plasmids
		GenScript Custom cDNA clone Services for MARVELD1

Primers:		OriGene genome-wide validated SYBR primer pairs: MARVELD1 Browse OriGene validated miRNA SYBR primer pairs
		SABiosciences RT² qPCR Primer Assay for MARVELD1: PPH14442A

REFSEQ mRNAs for MARVELD1 gene:

NM_031484.3

Additional cDNA sequence:

AF325259.1 AK025996.1 AK027093.1 CR596454.1

8 DOTS entries:

DT.217694 DT.97847681 DT.92434813 DT.100701883 DT.95276563 DT.95276564 DT.95276574 DT.95276565

24/270 AceView cDNA sequences (see all 270):

CR596454 AA339165 F03684 BX090352 BQ685706 CA428800 AI347102 BU789782

BU623551 AI452529 BQ722260 AA774589 BU195874 BU617019 W81344 AW419078

BM756030 W57786 BM971765 Z38491 BU753022 CD679436 BM982618 BU844661

highest scoring ESTs for MARVELD1:

AA151788 AA151833 AA293013 AA328333 AA341490 AA379476 AA402557 AA426289 AA446921 AA476612

Unigene Cluster for MARVELD1:

MARVEL domain containing 1

Hs.209614 [show with all ESTs]

Unigene Representative Sequence: NM_031484

GeneLoc Exon Structure

3 Alternative Splicing Database (ASD) splice patterns (SP) for MARVELD1

ExUns:	1	^	2a	·	2b	^	3a	·	3b	^	4a	·	4b
SP1:			-
SP2:							-
SP3:

About this scheme

ECgene alternative splicing isoforms for MARVELD1
5 Ensembl transcripts including schematic representations, and UCSC links where relevant:

ENST00000449755 ENST00000434038 ENST00000451097 ENST00000285605(uc001koj.3)

ENST00000421644

(Experimental results according to ¹GeneNote and GNF BioGPS,
probe sets-to-genes annotations according to ²GeneAnnot , ³GeneTide , Sets of similar genes according to GeneDecks, Electronic Northern calculations according to data from UniGene (Build 228 Homo sapiens), SAGE tags according to CGAP, plus additional links to SOURCE, and/or GNF BioGPS, and/or EXPOLDB, and/or UniProtKB,
Primers from OriGene and/or SABiosciences )
About This Section

MARVELD1 expression in normal and diseased human tissues

¹ / ² / ³

5 probe-sets matching MARVELD1 gene

Affymetrix probe-set	Array	# genes	Sensitivity	Specificity	Correlation	Length	Gb_Accession	Consensus	Uniqueness	Score	Rank
Affymetrix probe-set	Array	GeneAnnot data			GeneNote data		GeneTide data
57233_at²^{, 3}	U95-B	1	1.00	1.00	1.00	1.00	AI557563	0.60	1.00	0.82	1
223095_at²^{, 3}	U133-B	1	1.00	1.00	--	--	BC004995	0.80	1.00	0.91	1
234917_at²^{, 3}	U133-B	1	0.46	1.00	--	--	AK027093	0.80	1.00	0.91	1
223095_at²	U133Plus2	1	1.00	1.00	--	--	--	--	--	--	--
234917_at²	U133Plus2	1	0.46	1.00	--	--	--	--	--	--	--

About this table

MARVELD1 for expression About GeneDecksing

Data from Genenote (Publications) and GNF BioGPS
About these images

MARVELD1 gene electronic northern expression

About these images

CGAP SAGE TAG: GAGCCTCATC

SOURCE GeneReport for Unigene cluster: Hs.209614

UniProtKB/Swiss-Prot: MALD1_HUMAN, Q9BSK0

Tissue specificity: Widely expressed in normal tissues

Primers:		OriGene genome-wide validated SYBR primer pairs: MARVELD1 Browse OriGene validated miRNA SYBR primer pairs
		SABiosciences RT² qPCR Primer Assay for MARVELD1: PPH14442A

SABiosciences Custom PCR Arrays for MARVELD1

(Orthologs according to ^1,2HomoloGene (²older version, for species not in ¹newer version), ³euGenes, ⁴SGD and/or ⁵MGI Dec 24 2010, with possible further links to Flybase and/or WormBase, Gene Trees according to Ensembl)
About This Section

Orthologs for MARVELD1 gene from 2 species

Organism	Gene	Locus	Description	Human Similarity	NCBI accessions
mouse (Mus musculus)	Marveld1⁵	19 (35.84 cM)	MARVEL (membrane-associating) domain containing 1	--	BC054384
rainbow trout (Oncorhynchus mykiss)	Omy.5523²	--	Oncorhynchus mykiss transcribed sequence with weak more	75.34(n)	BX302064.1

About this table Species with no ortholog for MARVELD1

ENSEMBL Gene Tree for MARVELD1 (if available)

(Paralogs according to ¹HomoloGene
and ²Ensembl, Pseudogenes according to ³Pseudogene.org)
About This Section

(SNPs according to the ¹NCBI SNP Database, ²Ensembl, ³PupaSUITE, and UniProtKB, Linkage Disequilibrium by HapMap, Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Resequencing Primers from Qiagen)
About This Section

SNP ID	Valid	Chr 10 pos	Sequence	Recs	AA Chg	Type	More	Recs	Allele freq	Pop	Total sample	More
Genomic Data				Transcription Related Data				Allele Frequencies
			--	--	--		--	--
rs8231^1,2	C,F,	99477689(-)	`TGAGTC/GCTGGT`	1	--	ut3¹ ese³		2		MN WA	234
rs1043854^1,2	--	99477304(+)	`TTTTTC/TCTTTT`	1	--	ut3¹		0	--	--	--	--
rs1043855^1,2	A	99477305(+)	`TTTTCC/TTTTTT`	1	--	ut3¹		0	--	--	--	--
rs1043904^1,2	--	99477713(+)	`CTACTC/TTGGGC`	1	--	ut3¹ ese³		1		MN	184
rs1043907^1,2	--	99477766(+)	`GATTTC/TTACAG`	1	--	ut3¹ ese³		1		MN	184
rs4919133^1,2	C,F,A,	99476017(+)	`gctcaA/Ctgcag`	1	--	int¹		3		NA WA	76
rs6584148^1,2	C,F,A,H,	99472221(+)	`TgaccT/Cggctt`	1	--	ng5¹		8		NS EA NA WA	498
rs6584149^1,2	C,F,A,H,	99478081(+)	`GGAGCC/TTGGGC`	1	--	ng3¹		15		NS EA NA WA	1056
rs7896853^1,2	C,F,A,	99476915(+)	`GGTGCC/TTTGGA`	1	--	int¹		15		NS EA NA WA	998
rs11189341^1,2	--	99473379(+)	`agacaG/Tccgag`	1	--	ng5¹		0	--	--	--	--

About this table

HapMap Linkage Disequilibrium images for MARVELD1 (up to first 250kb)
Structural Variations (Copy Number Variations, Insertions/Deletions, Inversions)
Database of Genomic Variants (DGV) variations for MARVELD1: --

QIAGEN SeqTarget long-range PCR primers for resequencing MARVELD1

(in which this Gene is Involved, According to OMIM, UniProtKB, PharmGKB, Genatlas, GeneTests, Blood group antigen gene mutations by BGMUT, LSDB, HGMD, GAD, HuGE Navigator, and/or TGDB.)
About This Section

(Possibly Related Articles in Doctor's Guide)
About This Section

(in PubMed. Associations of this gene to articles via ¹Entrez Gene, ²UniProtKB/Swiss-Prot, ³HGNC, ⁴GAD, ⁵PharmGKB, ⁶UniProtKB/TrEMBL, and/or ⁷Novoseek)
About This Section

5 PubMed articles for MARVELD1 gene integrated from 7 sources:
(articles sorted by number of sources associating them with MARVELD1)

Identification and characterization of MARVELD1, a novel nuclear protein that is down-regulated in multiple cancers and silenced by DNA methylation. (PubMed id 19364627)^{1, 2} Wang S.... Wu H. (2009)
The DNA sequence and comparative analysis of human chromosome 10. (PubMed id 15164054)^{1, 2} Deloukas P.... Rogers J. (2004)
The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)^{1, 2} Gerhard D.S....Malek J. (2004)
Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. (PubMed id 12477932)^{1, 3} Strausberg R.L....Marra M.A. (2002)
Normalization and subtraction: two approaches to facilitate gene discovery. (PubMed id 8889548)¹ Bonaldo M.F.... Soares M.B. (1996)

(in PubMed, OMIM, and NCBI Bookshelf)
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		AND	OR
Aliases
Free Text

Query String		PubMed OMIM NCBI Bookshelf
	(Note: In FireFox, select the above section and copy using Ctrl-C)

(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, Kegg, and/or H-InvDB)
About This Section

Entrez Gene: 83742	HGNC: 28674	AceView: MARVELD1	Ensembl:ENSG00000155254	euGenes: HUgn83742
ECgene: MARVELD1	H-InvDB: MARVELD1

(According to HUGE)
About This Section

(According to ATLAS, HORDE, IMGT, MTDB, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
About This Section

Intellectual Property for MARVELD1 gene
(Patent information from GeneIP, Licensable technologies from WIS Yeda, Salk, Tufts, IP news from XenneX, Inc.)

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Patent Information for MARVELD1 gene:
Search GeneIP for patents involving MARVELD1

GeneCards and IP:
Japan Patent Office Licenses GeneCards European Patent Office Licenses GeneCards Improving the IP Search

(Antibodies, recombinant proteins, and assays by Millipore, Sigma-Aldrich, R&D Systems, Qiagen, GenScript, Cell Signaling Technology, SABiosciences, Novus Biologicals, Epitomics, ProSpec, Uscn,
Clones available from Millipore, Sigma-Aldrich, OriGene, GenScript, Sino Biological, PCR Arrays from SABiosciences, Drugs and/or compounds by Sigma-Aldrich, Tocris Bioscience, and/or Enzo Life Sciences)
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	QIAGEN Custom miScript Target Protector blocks miRNA-binding site of MARVELD1
	SeqTarget long-range PCR primers for resequencing MARVELD1
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MARVELD1 Gene

protein-coding GIFtS: 41

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MARVELD1 Gene

protein-coding GIFtS: 41

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Descriptions
for MARVELD1 gene

Summaries
for MARVELD1 gene

Genomic Views
for MARVELD1 gene

Proteins
for MARVELD1 gene

Protein Domains/
Families
for MARVELD1 gene

Gene Function
for MARVELD1 gene

Pathways & Interactions
for MARVELD1 gene

Drugs & Compounds
for MARVELD1 gene

Transcripts
for MARVELD1 gene

Expression
for MARVELD1 gene

Orthologs
for MARVELD1 gene

Paralogs
for MARVELD1 gene

Genomic Variants
for MARVELD1 gene

Disorders & Mutations
for MARVELD1 gene

Medical News
for MARVELD1 gene

Publications
for MARVELD1 gene

Intellectual Property
for MARVELD1 gene

Products
for MARVELD1 gene