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TSKU Gene

protein-coding   GIFtS: 47

GC11P072791
tsukushi small leucine rich proteoglycan homolog (Xenopus laevis)
(Previous names: leucine rich repeat containing 54, tsukushin )
Symbol approved by the HUGO Gene Nomenclature Committee (HGNC) database
(Previous symbol: LRRC54)
Products    

(According to 1HGNC, 2Entrez Gene,
3UniProtKB/Swiss-Prot, 4UniProtKB/TrEMBL, 5OMIM, 6GeneLoc , and/or 7Ensembl, 8miRBase)
About This Section

Aliases & Descriptions
tsukushi small leucine rich proteoglycan homolog (Xenopus laevis)1 2     LRRC542 3
TSK1 2 3 5     E2-induced gene 4 protein2 3
E2IG41 2 3 5     tsukushi homolog2
tsukushin1 2     tsukushi, small leucine rich proteoglycan2
leucine rich repeat containing 541 2     Tsukushi3
Leucine-rich repeat-containing protein 542 3     

External Ids:    HGNC: 288501   Entrez Gene: 259872   Ensembl: ENSG000001827047   UniProtKB: Q8WUA83   

Export aliases for TSKU gene to outside databases

Previous GC identifers: GC11P076171 GC11P076494


(According to Entrez Gene, Tocris Bioscience, Wikipedia's Gene Wiki,
UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL)
About This Section

  --

(According to GeneLoc and/or HGNC, and/or
Entrez Gene (NCBI build 37),
and/or miRBase,
Genomic Views according to UCSC (hg19) and Ensembl (release 60), Regulatory elements and Epigenetics data according to Qiagen and/or SABiosciences)
About This Section

Regulatory elements:
   Search SABiosciences Regulatory transcription factor binding sites for TSKU
         Other transcription factors

   Search SABiosciences Chromatin IP Primers for TSKU

Epigenetics:
QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays for TSKU 


Genomic Location:
Genomic View: UCSC Golden Path with GeneCards custom track

Entrez Gene cytogenetic band: 11q13.5   Ensembl cytogenetic band:  11q13.5   HGNC cytogenetic band: 11q13.5

TSKU Gene in genomic location: bands according to Ensembl, locations according to (and/or Entrez Gene and/or Ensembl if different)
TSKU gene location

GeneLoc gene densities for chromosome 11         GeneLoc Exon Structure

GeneLoc location for GC11P072791:  view genomic region     (about GC identifiers)

Start:
72,791,984 bp from pter
End:
72,806,897 bp from pter
Size:
14,914 bases
Orientation:
plus strand

1 alternative location:
Chr11+ 76,494,285-76,509,197     
RefSeq DNA sequence:
NC_000011.9  NT_167190.1  

(According to 1UniProtKB, neXtProt, and/or Ensembl, Phosphorylation sites according to 2Phosphosite, RefSeq according to NCBI, PDB rendering according to OCA and/or Proteopedia, Recombinant Proteins from Millipore, Sigma-Aldrich, R&D Systems, GenScript, Enzo Life Sciences, OriGene, Novus Biologicals, Sino Biological, and/or ProSpec,
Biochemical Assays by Millipore, Sigma-Aldrich, R&D Systems, OriGene, GenScript, Cell Signaling Technology, Enzo Life Sciences, and/or Uscn, Ontologies according to Gene Ontology Consortium 01 Dec 2010 and Entrez Gene, Antibodies by Millipore, Sigma-Aldrich, R&D Systems, GenScript, Cell Signaling Technology, OriGene, Novus Biologicals, and/or Epitomics)
About This Section

UniProtKB/Swiss-Prot: TSK_HUMAN, Q8WUA8 (See protein sequence)
Recommended Name: Tsukushin precursor  
Size: 353 amino acids; 37807 Da
Subcellular location: Secreted (Potential)
Secondary accessions: B3KQT7 Q6UXK1 Q9UG10 Q9UJX9

Explore the universe of human proteins at neXtProt for TSKU: NX_Q8WUA8 

REFSEQ proteins: NP_056331.2  

ENSEMBL proteins: 
ENSP00000332668 ENSP00000387819 


Human Recombinant Proteins 
Browse Purified and Recombinant Proteins at Millipore
Browse Human Recombinant Proteins at Sigma-Aldrich
R&D Systems Recombinant & Natural Proteins for TSKU (Tsukushi/TSK)
Browse recombinant and purified proteins available from Enzo Life Sciences
Browse OriGene full length recombinant human proteins expressed in human HEK293 cells
OriGene Protein Over-expression Lysate: TSKU
GenScript Custom Purified and Recombinant Proteins Services for TSKU 
Novus Biologicals Proteins for TSKU
Novus Biologicals Lysates for TSKU
Browse Sino Biological Recombinant Proteins
Browse ProSpec Recombinant Proteins

1 Gene Ontology (GO) cellular component term (GO ID links to tree view):

GO IDQualified GO termEvidencePubMed IDs
GO:0005576 extracellular region IEA--
About this table

TSKU for ontologies           About GeneDecksing



Antibodies for TSKU: 
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Browse antibodies at Epitomics

Assays for TSKU: 
Browse Kits and Assays available from Millipore
Browse ELISAs at Sigma-Aldrich
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Browse OriGene Fluorogenic Cell Assay Kits 
Browse R&D Systems for biochemical assays
GenScript Custom Assay Services for TSKU 
Browse Enzo Life Sciences for kits & assays
Browse ELISAs and CLIAs at Uscn 


(According to InterPro, ProtoNet, UniProtKB, and/or BLOCKS, Sets of similar genes according to GeneDecks)
About This Section

TSKU for domains           About GeneDecksing

3 InterPro domains/families:
 IPR001611 Leu-rich_rpt
 IPR003591 Leu-rich_rpt_typical-subtyp
 IPR000372 LRR-contain_N

Graphical View of Domain Structure for InterPro Entry Q8WUA8

ProtoNet protein and cluster: Q8WUA8

1 Blocks protein family: IPB000372 Cysteine-rich flanking region

UniProtKB/Swiss-Prot: TSK_HUMAN, Q8WUA8
Similarity: Contains 9 LRR (leucine-rich) repeats


(According to UniProtKB, IUBMB,and/or Genatlas, Animal models from MGI Dec 24 2010,
shRNA from OriGene, Sigma-Aldrich, RNAi from Millipore, siRNAs from Sigma-Aldrich, OriGene, Qiagen, Super-pooled esiRNAs from Sigma-Aldrich, microRNA from Sigma-Aldrich, Qiagen, SABiosciences, Clones from Millipore, Sigma-Aldrich, OriGene, GenScript, Sino Biological, Cell Lines from GenScript, Ontologies according to Gene Ontology Consortium 01 Dec 2010 via Entrez Gene.)
About This Section

UniProtKB/Swiss-Prot: TSK_HUMAN, Q8WUA8
Induction: By 17-beta-estradiol

Inhib.
RNA:
    
Browse for Gene Knock-down Tools from Millipore
Browse Nano Scale siRNA at Sigma-Aldrich
Sigma-Aldrich esiRNA and siRNA for TSKU
Sigma-Aldrich shRNA for TSKU
OriGene 29mer shRNA kits in GFP-retroviral vector: TSKU
OriGene shRNA RFP: TSKU
OriGene basic RS shRNA: TSKU
OriGene siRNA: TSKU
QIAGEN FlexiTube/FlexiPlate siRNA for gene silencing of TSKU 
miRNA:Sigma-Aldrich microRNA Mimics for TSKU
Browse microRNA target validation systems at Sigma-Aldrich 
QIAGEN Custom miScript Target Protector blocks miRNA-binding site of TSKU 
5 SABiosciences Assays for microRNAs that regulate TSKU:
hsa-miR-492 hsa-miR-622 hsa-miR-1271 hsa-miR-663b hsa-miR-96

Gene
Editing:
Browse CompoZr Knockout ZFN at Sigma-Aldrich 

Clones:Browse Clones for the Expression of Recombinant Proteins Available from Millipore
Browse iPSC Reprogramming Factors at Sigma-Aldrich
OriGene GFP tagged cDNA clones in CMV expression vector: TSKU
OriGene Myc/DDK tagged cDNA clones in CMV expression vector: TSKU
OriGene untagged cDNA clones in CMV expression vector (see all 2): TSKU
OriGene 3'-UTR clone: TSKU 
Browse MicroRNA Expression Plasmids 
GenScript cDNA clone with any tag delivered in your preferred vector: TSKU 
Sino Biological Human cDNA Clone for TSKU

Cell
Lines:
  
GenScript Custom overexpressing Cell Line Services for TSKU 

1 Gene Ontology (GO) molecular function term (GO ID links to tree view):

GO IDQualified GO termEvidencePubMed IDs
GO:0005515 protein binding IEA--
About this table

TSKU for ontologies           About GeneDecksing


(Pathways according to Millipore, Cell Signaling Technology, Sigma-Aldrich, KEGG and/or UniProtKB, Sets of similar genes according to GeneDecks, PCR Arrays from SABiosciences, Proteins Network according to SABiosciences, Sigma-Aldrich, Interactions according to 1UniProtKB, 2MINT, and/or 3STRING, with links to IntAct and Ensembl, Ontologies according to Gene Ontology Consortium 01 Dec 2010 via Entrez Gene).
About This Section

    SABiosciences Custom Pathway-Focused PCR Arrays for TSKU 

    SABiosciences Gene Network CentralTM Interacting Genes and Proteins Network for TSKU

1 Interacting protein for TSKU (Q8WUA82) via UniProtKB, MINT, and/or STRING
InteractantInteraction Details
GeneCardExternal ID(s)
SKILP127572MINT-61667
About this table

(Chemical Compounds according to UniProtKB, Enzo Life Sciences, Sigma-Aldrich, Tocris Bioscience, and/or Novoseek and Drugs according to Enzo Life Sciences and/or PharmGKB)
About This Section
Browse Small Molecules at Sigma-Aldrich
Browse drugs & compounds from Enzo Life Sciences

Browse Tocris compounds for TSKU


(GenBank/EMBL/DDBJ Accessions according to
Unigene (Build 228 Homo sapiens; Dec 8 2010) or GenBank,
RefSeq according to Entrez Gene,
DOTS (version 10), and/or AceView, transcript ids from Ensembl with links to UCSC,
non coding RNAs according to RNAdb,
ESTs according to GeneTide,
exon structure from GeneLoc, alternative splicing isoforms according to ASD and/or ECgene,
RNAi Products from Millipore,
siRNAs from Sigma-Aldrich, OriGene, Qiagen, Super-pooled esiRNAs from Sigma-Aldrich,
shRNA from Sigma-Aldrich, OriGene, microRNA from Sigma-Aldrich, Qiagen, SABiosciences,
Tagged/untagged cDNA clones from OriGene, Sigma-Aldrich, GenScript, Primers from OriGene and/or SABiosciences)
About This Section

Inhib.
RNA:
     
Browse for Gene Knock-down Tools from Millipore
Browse Nano Scale siRNA at Sigma-Aldrich
Sigma-Aldrich esiRNA and siRNA for TSKU
Sigma-Aldrich shRNA for TSKU
OriGene 29mer shRNA kits in GFP-retroviral vector: TSKU
OriGene shRNA RFP: TSKU
OriGene basic RS shRNA: TSKU
OriGene siRNA: TSKU
QIAGEN FlexiTube/FlexiPlate siRNA for gene silencing of TSKU 
miRNA: Sigma-Aldrich microRNA Mimics for TSKU
Browse microRNA target validation systems at Sigma-Aldrich 
QIAGEN Custom miScript Target Protector blocks miRNA-binding site of TSKU 
5 SABiosciences Assays for microRNAs that regulate TSKU:
hsa-miR-492 hsa-miR-622 hsa-miR-1271 hsa-miR-663b hsa-miR-96
Clones: Browse FLAG tag genes at Sigma-Aldrich
OriGene GFP tagged cDNA clones in CMV expression vector: TSKU
OriGene Myc/DDK tagged cDNA clones in CMV expression vector: TSKU
OriGene untagged cDNA clones in CMV expression vector (see all 2): TSKU
OriGene 3'-UTR Clone: TSKU 
Browse OriGene MicroRNA Expression Plasmids 
GenScript cDNA clone with any tag delivered in your preferred vector: TSKU 
Primers: OriGene genome-wide validated SYBR primer pairs: TSKU
Browse OriGene validated miRNA SYBR primer pairs 
SABiosciences RT2 qPCR Primer Assay for TSKU: PPH07614A

REFSEQ mRNAs for TSKU gene: 

NM_015516.3  

Additional cDNA sequence: 

AF191019.1 AK075476.1 AK091475.1 AK126536.1 AK313405.1 AL110276.1 AY358317.1 BC020975.1 
BT007440.1 

10 DOTS entries:

DT.417669  DT.100777424  DT.100754481  DT.120701703  DT.100777423  DT.91768837  DT.91768832  DT.120701695 
DT.120701697  DT.40215150 

24/258 AceView cDNA sequences (see all 258):

BG697318 BF724678 AI884558 BE336901 AI269100 BF724254 BU526767 AI074139 
BE786245 BU846213 AA558470 BF725906 BM723751 CA489754 BG056788 AI074399 
CB242473 AI613032 BX324381 AA305745 BC020975 AA335297 AL110276 BG327574 

highest scoring ESTs for TSKU:

AI702079 AA009451 AA009735 AA045801 AA045868 AA088181 AA299901 AA305745 AA335281 AA335297 

Unigene Cluster for TSKU:

Tsukushi small leucine rich proteoglycan homolog (Xenopus laevis)
Hs.8361  [show with all ESTs]
Unigene Representative Sequence: AK126536


GeneLoc Exon Structure

2 Ensembl transcripts including schematic representations, and UCSC links where relevant:
ENST00000333090(uc001oxt.2) ENST00000439807

(Experimental results according to 1GeneNote and GNF BioGPS,
probe sets-to-genes annotations according to 2GeneAnnot , 3GeneTide , Sets of similar genes according to GeneDecks, Electronic Northern calculations according to data from UniGene (Build 228 Homo sapiens), SAGE tags according to CGAP, plus additional links to SOURCE, and/or GNF BioGPS, and/or EXPOLDB, and/or UniProtKB,
Primers from OriGene and/or SABiosciences )
About This Section

TSKU expression in normal and diseased human tissues

1  / 2  / 3

3 probe-sets matching TSKU gene

Affymetrix
probe-set
Array  GeneAnnot data GeneNote data GeneTide data
# genes Sensitivity Specificity Correlation Length Gb_Accession Consensus Uniqueness Score Rank
46313_at2, 3 U95-B 1 1.00 1.00 1.00 1.00 AI702079 1.00 1.00 1.00 1
218245_at2, 3 U133-A 1 1.00 1.00 -- -- NM_015516 0.60 1.00 0.82 1
218245_at2 U133Plus2 1 1.00 1.00 -- -- -- -- -- -- --
About this table

TSKU for expression           About GeneDecksing

Data from Genenote  (Publications) and GNF BioGPS
    About these images
TSKU gene expression
TSKU gene electronic northern expression
TSKU gene sage expression
About these images

CGAP SAGE TAG: GGCATCAGGG

SOURCE GeneReport for Unigene cluster: Hs.8361
Primers: OriGene genome-wide validated SYBR primer pairs: TSKU
Browse OriGene validated miRNA SYBR primer pairs 
SABiosciences RT2 qPCR Primer Assay for TSKU: PPH07614A
    SABiosciences Custom PCR Arrays for TSKU 


(Orthologs according to 1,2HomoloGene (2older version, for species not in 1newer version), 3euGenes, 4SGD and/or 5MGI Dec 24 2010, with possible further links to Flybase and/or WormBase, Gene Trees according to Ensembl)
About This Section

Orthologs for TSKU gene from 5/8 species (see all 8)
Organism Gene Locus Description Human
Similarity
NCBI accessions
dog
(Canis familiaris)
TSKU1   -- tsukushin 86.5(n)
87.25(a)
485178  XM_542296.2  XP_542296.2 
chimpanzee
(Pan troglodytes)
TSKU1   -- tsukushin 98.75(n)
97.7(a)
451436  XM_508654.2  XP_508654.2 
cow
(Bos taurus)
TSKU1   -- tsukushin 86.69(n)
84.99(a)
529661  XM_866913.2  XP_872006.1 
rat
(Rattus norvegicus)
Tsku1   -- tsukushin 81.56(n)
85.59(a)
308843  NM_001009965.2  NP_001009965.1 
mouse
(Mus musculus)
Tsku1 , 5 7 (53.66 cM)5
tsukushin1, 5 82.57(n)1
85.63(a)1
2441521  NM_001024619.21  NP_001019790.11 
 AB2141845  AK0354615  (see all 12)
About this table        Species with no ortholog for TSKU

ENSEMBL Gene Tree for TSKU (if available)

(Paralogs according to 1HomoloGene
and 2Ensembl, Pseudogenes according to 3Pseudogene.org)
About This Section
  --

(SNPs according to the 1NCBI SNP Database, 2Ensembl, 3PupaSUITE, and UniProtKB, Linkage Disequilibrium by HapMap, Structural Variations(CNVs/InDels/Inversions) from the Database of Genomic Variants, Resequencing Primers from Qiagen)
About This Section

10/165 NCBI SNPs in TSKU are shown (see all 165)
Genomic DataTranscription Related DataAllele Frequencies
SNP IDValidChr 11 posSequenceRecsAA
Chg
TypeMoreRecsAllele
freq
PopTotal
sample
More
----------
rs79121,2
C,F,A,76508843(+) CAGGGT/CAGCAT 1 -- ut318Minor allele frequency- C:0.16MN NS EA NA 866
rs11496071,2
C,F,A,76492446(-) GACTGG/TAAAGC 1 -- ng51 tfbs33Minor allele frequency- T:0.31NA EA WA 210
rs11496081,2
76494928(-) CTCTCA/GGTGCG 1 -- int10--------
rs11496091,2
C,A,76495113(-) ATCCCA/GTCGCA 1 -- int10--------
rs11496101,2
C,F,76495497(-) GAGGCT/ACCCAC 1 -- int11Minor allele frequency- A:0.22NA 72
rs11496111,2
C,F,O,A,76495879(-) TGAGCG/AGTGGG 1 -- int121Minor allele frequency- A:0.39EA MN NA NS WA 3048
rs11496121,2
C,F,A,76496736(+) GGGGAT/CTTGAG 1 -- int13Minor allele frequency- C:0.24MN NA EA 344
rs11496131,2
C,F,A,H,76497432(-) CATTTC/GATTCA 1 -- int1111Minor allele frequency- G:0.49NS MN EA PA EU CA WA NA 7332
rs11496141,2
C,F,76497563(-) TAAAAC/TCCATC 1 -- int14Minor allele frequency- T:0.22MN NA EA WA 394
rs11496151,2
C,F,76498133(-) ctaacG/Aggtca 1 -- int110Minor allele frequency- A:0.26NS EA NA 918
About this table

HapMap Linkage Disequilibrium images for TSKU (up to first 250kb)
Structural Variations (Copy Number Variations, Insertions/Deletions, Inversions)
      Database of Genomic Variants (DGV) variations for TSKU: --

QIAGEN SeqTarget long-range PCR primers for resequencing TSKU 

(in which this Gene is Involved, According to OMIM, UniProtKB, PharmGKB, Genatlas, GeneTests, Blood group antigen gene mutations by BGMUT, LSDB, HGMD, GAD, HuGE Navigator, and/or TGDB.)
About This Section

TSKU for disorders           About GeneDecksing

OMIM: 608015


Export disorders and mutations for TSKU gene to outside databases

(Possibly Related Articles in Doctor's Guide)
About This Section
  --

(in PubMed. Associations of this gene to articles via 1Entrez Gene, 2UniProtKB/Swiss-Prot, 3HGNC, 4GAD, 5PharmGKB, 6UniProtKB/TrEMBL, and/or 7Novoseek)
About This Section

9 PubMed articles for TSKU gene integrated from 7 sources:
(articles sorted by number of sources associating them with TSKU)
  1. Effects of estrogen on global gene expression: identification of novel targets of estrogen action. (PubMed id 11085516)1, 2, 3 Charpentier A.H.... Aldaz C.M. (2000)
  2. Comparative analysis of gene regulation by the transc ription factor PPARalpha between mouse and human. (PubMed id 19710929)1, 3 Rakhshandehroo M....Kersten S. (2009)
  3. Signal sequence and keyword trap in silico for selection of full- length human cDNAs encoding secretion or membrane proteins from oligo- capped cDNA libraries. (PubMed id 16303743)1, 2 Otsuki T....Isogai T. (2005)
  4. The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). (PubMed id 15489334)1, 2 Gerhard D.S....Malek J. (2004)
  5. The secreted protein discovery initiative (SPDI), a large-scale effort to identify novel human secreted and transmembrane proteins: a bioinformatics assessment. (PubMed id 12975309)1, 2 Clark H.F.... Gray A.M. (2003)
  6. The full-ORF clone resource of the German cDNA consortium. (PubMed id 17974005)2 Bechtel S.... Schupp I. (2007)
  7. Functional proteomics mapping of a human signaling pathway. (PubMed id 15231748)1 Colland F....Gauthier J.M. (2004)
  8. Complete sequencing and characterization of 21,243 full-length human cDNAs. (PubMed id 14702039)1 Ota T.... Sugano S. (2004)
  9. Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. (PubMed id 12477932)1 Strausberg R.L....Marra M.A. (2002)

(in PubMed, OMIM, and NCBI Bookshelf)
About This Section
 ANDOR
Aliases
Free Text  

  Query String
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OMIM
NCBI Bookshelf
  (Note: In FireFox, select the above section and copy using Ctrl-C)

(According to Entrez Gene, HGNC, AceView, euGenes, Ensembl, miRBase, ECgene, Kegg, and/or H-InvDB)
About This Section
Entrez Gene: 25987 HGNC: 28850 AceView: TSK Ensembl:ENSG00000182704 euGenes: HUgn25987
ECgene: TSKU H-InvDB: TSKU

(According to HUGE)
About This Section
  --

(According to ATLAS, HORDE, IMGT, MTDB, LEIDEN, UniProtKB/Swiss-Prot, and/or UniProtKB/TrEMBL,
Wikipedia and/or GeneReviews via UniProtKB/Swiss-Prot)
About This Section
  --

(Patent information from GeneIP,
Licensable technologies from WIS Yeda, Salk, Tufts,
IP news from XenneX, Inc.)
About This Section
Patent Information for TSKU gene:
Search GeneIP for patents involving TSKU

GeneCards and IP:
Japan Patent Office Licenses GeneCards     European Patent Office Licenses GeneCards     Improving the IP Search



(Antibodies, recombinant proteins, and assays by Millipore, Sigma-Aldrich, R&D Systems, Qiagen, GenScript, Cell Signaling Technology, SABiosciences, Novus Biologicals, Epitomics, ProSpec, Uscn,
Clones available from Millipore, Sigma-Aldrich, OriGene, GenScript, Sino Biological, PCR Arrays from SABiosciences, Drugs and/or compounds by Sigma-Aldrich, Tocris Bioscience, and/or Enzo Life Sciences)
About This Section

 Millipore Custom Antibody & Bulk Services
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 Predesigned and custom siRNAs for TSKU Antibodies for TSKU
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 "Your Favorite Gene" Pathways and Molecular Interaction Networks Browse Nano Scale siRNA at Sigma-Aldrich
 Browse iPSC Reprogramming Factors at Sigma-Aldrich Browse small molecules at Sigma-Aldrich
 Browse FLAG tag genes at Sigma-Aldrich Browse ELISAs at Sigma-Aldrich
 Browse microRNA Target Validation Systems at Sigma-Aldrich Browse CompoZr Knockout ZFN at Sigma-Aldrich
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 miRNA Expression Plasmids

 
 
 QIAGEN Custom miScript Target Protector blocks miRNA-binding site of TSKU
 SeqTarget long-range PCR primers for resequencing TSKU
 QIAGEN PyroMark CpG Assay predesigned Pyrosequencing DNA Methylation assays for TSKU
 QIAGEN FlexiTube/FlexiPlate siRNA for gene silencing of TSKU
 GenScript Custom Purified and Recombinant Proteins Services for TSKU GenScript cDNA clones with any tag delivered in your preferred vector for TSKU
 GenScript Custom Assay Services for TSKU GenScript Custom Superior Antibodies Services for TSKU
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